Noted dog trainer
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Related clues
- Ivan ___, Russian physiologist noted for his study of conditioned reflexes in dogs; Nobel Prize for Physiology or Medicine (1904)
- Ivan, Russian physiologist who studied reflexes in dogs
- Ivan ---, Russian physiologist who studied conditioned reflexes in dogs
- 'Conditioned reflex' Nobelist
- 1904 Physiology Nobelist
Possible answers
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- A
- V
- L
- O
- V